The below publications are protected by copyright laws. Please email kellylabumanitoba @ to request copies.

Underlined names indicates work completed as a student or postdoc:


  • Clary, D., Stow, M.K., Vernouillet, A., & Kelly, D.M.. (Accepted pending revisions). Mirror-meditated responses of California scrub jays during a caching task and the mark task. Ethology
  • Kelly, D.M., Adolphe, J.L., Vernouillet, A., McCausland, J.A., Rankovic, A., & Verbrugghe, A. (2019). Motoric Self-Regulation by Sled Dogs and Pet Dogs and the Acute Effect of Carbohydrate Source in Sled Dogs. Animal Cognition. Full Article
  • Ranjbar Pouya, O., Byagowi, A., Kelly, D.M., & Moussavi, Z. (In Press). Comparisons of target localization abilities during physical and virtual rotating scenes by cognitively-intact and cognitively-impaired older adults. OBM Geriatrics
  • Kelly, D.M., Bisbing, T., & Magnotti, J. (2019). Use of medial axis for reorientation by the Clark's nutcracker (Nucifraga columbiana). Behavioural Processes, 158, 192-199 Full Article


  • Vernouillet, A., Stiles, L.R., McCausland, J.A., & Kelly, D.M.. (2018). Individual performance of motor self-regulation tasks is not correlated across tasks in pet dogs. Learning & Behaviour, Full Article
  • Kimura, K., Reichert, J.F., Pouya, O.R., Byagowi, R.R., Wang, X., Kelly, D.M., & Moussavi, Z. (2018). Simultaneous localization and mapping (SLAM) for route reversal learning. OBM Geriatrics, 2, doi:10.21926/obm.geriatr.1803007 Full Article
  • Kelly, D.M., Cheng, K., Balda, R.P., & Kamil, A.C. (2018). The effects of sun compass error on spatial search by Clark’s nutcrackers. Integrative Zoology, doi: 10.1111/1749-4877.12302. Abstract
  • Vernouillet, A.. (2018). Social intelligence hypothesis. In Encyclopedia of Animal Cognition and Behavior. Vonk, J., & Shackleford, T. D. Springer. Full Article
  • Stow, M.K., Vernouillet, A., & Kelly, D.M.. (2018). Neophobia does not account for motoric self-regulation performance as measured during the detour-reaching cylinder task. Animal Cognition, 21, 565-574. Full Article
  • Ranjbar Pouya, O., Kelly, D.M., & Moussavi, Z. (2018). Predicting cognitive status of older adults using directional accuracy in explicit timing tasks. Journal of Medical and Biological Engineering, Full Article
  • Wright, A.A., Kelly, D.M., & Katz, J.S. (2018). Comparing cognition by integrating concept learning, proactive interference, and list memory. Learning & Behavior, 46, 107-123. Full Article
  • Qadri, M.A., Leonard, K., Cook, R.G., & Kelly, D.M. (2018). Examination of long-term memorization capacity in the Clark’s nutcracker (Nucifraga columbiana). Psychonomic Bulletin & Review. Full Article
  • Leonard, K., Tian, N., Ivanco, T.L., & Kelly, D.M. (2018). Experience with Featural Cue Reliability Influences Featural and Geometric Cue Use by Mice (Mus musculus). Journal of Comparative Psychology, 132, 106-114. Abstract


  • Siemens, M. & Kelly, D.M. (2017). Sex differences and the effect of instruction on reorientation abilities by humans. Memory & Cognition, 46(4), 556-576. doi: 10.3758/s13421-017-0783-3 Full Article
  • Kimura, K., Reichert, J., Olson, A., Ranjbar Pouya, O., Moussavi, Z., & Kelly, D.M.. (2017). Orientation in virtual reality does not fully measure up to the real-world. DOI:10.1038/s41598-017-18289-8 Scientific Reports, 7.Full Article
  • Wright, A.A., Katz, J.S., & Kelly, D.M. (2017). Same/Different Learning. Encyclopedia of Animal Cognition and Behavior. October 24, 2017.
  • Wright A.A. & Kelly D.M. (2017). Comparative approaches to same/different abstract-concept learning Learning and Behaviour, 45(4), 323-324. doi: 10.3758/s13420-017-0271-4. Full Article
  • Magnotti, J.F., Wright, A.A., Katz J.S., Leonard K., Vernouillet, A., & Kelly D.M. (2017). Corvids Outperform Pigeons and Primates in Learning a Basic Concept. Psychological Science, 28(4), 437-444. DOI:10.1177/09567976/616685871. Full Article
  • Magnotti, J.F., Wright, A.A., Leonard, K., Katz, J.S., & Kelly, D.M. (2017). Abstract-concept learning in Black-billed magpies (Pica hudsonia). Psychonomic Bulletin & Review, 24(2), 431-435. doi: 10.3758/s13423-016-1132-4. Full Article
  • Ranjbar Pouya, O., Byagowi, A., Lissitsn, Y., Kelly, D.M., & Moussavi, Z. (2017). Introducing a new age- and cognition- sensitive measurement for assessing egocentric spatial orientation using a landmark-less virtual reality navigational task. Quarterly Journal of Experimental Psychology, 70, 1406-1419. DOI:10.1080/17470218.2016.1187181. Abstract


  • Clary, D., & Kelly D.M. (2016). Clark's Nutcrackers (Nucifraga columbiana) Flexibly Adapt Caching Behaviour to a Cooperative Context. Frontiers in Psychology. 7:1643. doi: 10.3389/fpsyg.2016.01643. Full Article
  • Vernouillet, A., Anderson, J., Clary, D., Kelly, D.M. (2016). Inhibition in Clark's nutcrackers (Nucifraga columbiana): results of a detour-reaching test. Animal Cognition, 19(3), 661-665. doi: 10.1007/s10071-016-0952-y Full Article
  • Wright, A.A., Magnotti, J.F., Katz, J.S., Leonard, K., & Kelly, D.M. (2016). Concept learning set-size functions for Clark's nutcrackers. Journal of the Experimental Analysis of Behavior. 105(1), 76-84. doi: 10.1002/jeab.174 Abstract


  • Magnotti,J.F.,Katz,J.S.,Wright,A.A., & Kelly,D.M.(2015). Superior abstract-concept learning by Clark’s nutcrackers (Nucifraga columbiana). Biology Letters, 11(5). doi: 10.1098/rsbl.2015.0148 Full Article
  • Reichert, J.F., & Kelly, D.M. (2015). How Clark's nutcrackers (Nucifraga columbiana) weigh geometric cues depends on their previous experience. Animal Cognition, 18(4), 953-968. DOI: 10.1007/s10071-015-0866-0. Full Article


  • Clary, D., Cheys, A., & Kelly, D.M. (2014). Pattern of visuospatial lateralization in two corvid species, black-billed magpies and Clark's nutcrackers. Behavioural Processes, 107, 94-98. Abstract
  • Kelly, D.M., & Cheng, K. (2014). Editorial: A synthetic approach to comparative cognition. Behavioural Processes, 102, 1-2. Abstract
  • Lambinet, V., Wilzeck, C., & Kelly, D.M. (2014). Size doesn't matter, but features do: Clark's nutcrackers (Nucifraga columbiana) weigh features more heavily than geometry in large and small enclosures. Behavioural Processes, 102, 3-11. Abstract


  • Snow, W.M., Stoesz, B.M., Kelly, D.M., & Albensi, B.C. (2013). Roles for NF-kB and Gene Targets of NF-kB in Synaptic Plasticity, Memory, and Navigation. Molecular Neurobiology, 49(2), 757-770 DOI 10.1007/s12035-013-8555-y Full Article
  • Ambosta, A.H., Reichert, J.F., & Kelly, D.M. (2013). Reorienting in virtual 3D environments: Do adult humans use principal axes, medial axes or local geometry? PLoS ONE, 8(11). Article.
  • Clary, D. & Kelly, D.M. (2013). Are Clark's nutcrackers (Nucifraga columbiana) able to discriminate knowledge states of human experimenters during an object-choice task? Evolutionary Psychology, 11, 1-37. Full Article
  • Sturz, B.R., & Kelly, D.M. (2013). Environment Size and the Use of Feature and Geometric Cues for Reorientation. Acta Psychologica, 142, 251-258. Abstract


  • Kelly, D.M., & Reichert, J.F. (2012). Angular amplitude matters: Exploring the functional relationship of geometric cue use by Clark's nutcrackers (Nucifraga columbiana). Behavioural Processes, 93, 62-70. Abstract
  • Sturz, B.R., Kilday, Z.A., Bodily, K.D., & Kelly, D.M. (2012). No evidence that consistent auditory cues facilitate learning of spatial relations among locations. Behavioural Processes, 90, 198-203. Abstract
  • Reichert, J.F. & Kelly, D.M. (2012). Discrimination of Geometric Angles by Adult Humans. Animal Cognition, 15, 271-284. doi: 10.1007/s10071-011-0452-z Full Article
  • Gibson, B., Wilkinson, M. & Kelly D.M. (2012). Let the pigeon drive the bus: pigeons can plan future routes in the real world. Animal Cognition, 15, 379-391. doi: 10.1007/s10071-011-0463-9 Full Article


  • Fairhurst, G.D., Frey, M.D., Reichert, J.F., Szelest, I., Kelly, D.M. & Bortolotti, G.R. (2011). Does Environmental Enrichment Reduce Stress? An Integrated Measure of Corticosterone from Feathers Provides a Novel Perspective. PLoS ONE, 6(3). Full Article
  • Kelly, D.M. & Durocher, S. (2011). Comparing Geometric Models for Orientation: Medial vs. Principal Axes. Communicative and Integrative Biology, 6, 710-712. Abstract
  • Kelly, D.M., Durocher, S., Chiandetti, C. & Vallortigara, G. (2011). A misunderstanding of principal and medial axes? Reply to Sturz & Bodily. Biology Letters, 7, 649-650. doi: 10.1098/rsbl.2011.0482 Full Article
  • Clary, D. & Kelly, D.M. (2011). Cache protection strategies of a non-social food-caching corvid, Clark’s nutcracker (Nucifraga columbiana). Animal Cognition, 14, 735-744. doi: 10.1007/s10071-011-0408-3 Full Article
  • Kelly, D.M., Chiandetti, C. & Vallortigara, G. (2011). Re-orienting in space: Do animals use global or local geometry strategies? Biology Letters, 7, 372-375. DOI: 10.1098/rsbl.2010.1024 Full Article
  • Reichert, J.F. & Kelly, D.M. (2011). Use of local and global geometry from object arrays by adult humans. Behavioural Processes, 86, 196-205.


  • Kelly, D.M. (2010). Features enhance encoding of geometry. Animal Cognition, 13, 453-462. Full Article
  • Rugani, R., Kelly, D.M., Szelest, I., Regolin, L. & Vallortigara, G. (2010). Is it only humans that count from left to right? Biology Letters, 6, 290-292. DOI: 10.1098/rsbl.2009.0960 Full Article
  • Gould, K.L., Kelly, D.M. & Kamil, A.C. (2010). What scatter-hoarding animals have taught us about small-scale navigation. Philosophical Transactions of the Royal Society B, 365, 901-914. DOI: 10.1098/rstb.2009.0214 Full Article
  • Sturz, B.R., Kelly, D.M. & Brown, M.F. (2010). Facilitation of learning spatial relations among locations by visual cues: Generality across spatial configurations. Animal Cognition, 13, 341-349. doi: 10.1007/s10071-009-0283-3 Full Article
  • Kelly, D.M., Kamil, A.C. & Cheng, K. (2010). Landmark use by Clark's nutcrackers (Nucifraga columbiana): Influence of disorientation and cue rotation on distance and direction estimates. Animal Cognition, 13, 175-188. doi: 10.1007/s10071-009-0256-6 Full Article
  • Dawson, M.R.W., Kelly, D.M., Spetch, M.L. & Dupuis, B. (2010). Using perceptrons to explore the reorientation task. Cognition, 114, 207-226. doi: 10.1016/j.cognition.2009.09.006 Abstract


  • Sturz, B.R. & Kelly, D.M. (2009). Encoding of relative enclosure size in a dynamic three-dimensional virtual environment by humans. Behavioural Processes, 82, 223-227. doi: 10.1016/j.beproc.2009.06.007 Abstract
  • Sturz, B.R., Brown, M.F. & Kelly, D.M. (2009). Facilitation of learning spatial relations among locations by visual cues: Implications for theoretical accounts of spatial learning. Psychonomic Bulletin & Review, 16, 306-312. doi: 10.3758/PBR.16.2.306 Full Article
  • Wilzeck, C., Prior, H. & Kelly, D.M. (2009). Geometry and landmark representation by pigeons: Evidence for species-differences in the hemispheric organization of spatial information processing? European Journal of Neuroscience, 4, 813-822. doi: 10.1111/j.1460-9568.2009.06626.x Abstract
  • Dawson, M.R.W., Dupuis, B., Spetch, M.L. & Kelly, D.M. (2009). Simple Artificial Neural Networks That Match Probability and Exploit and Explore When Confronting a Multiarmed Bandit. IEEE Transactions on Neural Networks, 20, 1368-1371. doi: 10.1109/TNN.2009.2025588 Abstract
  • Sturz, B.R., Bodily, K.D., Katz, J.S. & Kelly, D.M. (2009). Evidence against integration of spatial maps by humans (Homo sapiens). Animal Cognition, 12, 237-247. doi: 10.1007/s10071-008-0182-z Full Article


  • Kelly, D.M. & Bischof, W.F. (2008). Orienting in virtual environments: How are surface features and environmental geometry weighted in an orientation task? Cognition, 109, 89-104. doi: 10.1016/j.cognition.2008.07.012 Abstract
  • Dawson, M.R.W., Kelly, D.M, Spetch, M.L. & Dupuis, B. (2008). Learning about environmental geometry: A flaw in Miller and Shettleworth’s (2007) operant model. Journal of Experimental Psychology: Animal Behavior Processes, 34, 415-418. doi: 10.1037/0097-7403. Abstract
  • Kelly, D.M., Kippenbrock, S., Templeton, J. & Kamil, A.C. (2008). Use of a geometric rule or absolute vectors: Landmark use by Clark’s nutcrackers (Nucifraga columbiana). Brain Research Bulletin, 76, 293-299. doi: 10.1016/j.brainresbull.2008.02.008 Abstract



  • Cheng, K., Spetch, M.L., Kelly, D.M. & Bingman, V.P. (2006). Small-scale spatial cognition in pigeons. Behavioral Processes, 72, 115-127. DOI: 10.1016/j.beproc.2005.11.018 Abstract


  • Kelly, D.M. & Bischof, W.F. (2005). Re-orienting in images of a three-dimensional environment. Journal of Experimental Psychology: Human Perception and Performance, 31, 1391-1403. DOI: 10.1037/0096-1523.31.6.1391 Abstract


  • Kelly, D.M. & Spetch, M.L (2004b). Reorientation in a two-dimensional environment: II. Do pigeons encode the featural and geometric properties of a two-dimensional schematic of a room? Journal of Comparative Psychology, 118, 384-395. DOI: 10.1037/0735-7036.118.4.384 Abstract
  • Kelly, D.M. & Spetch, M.L (2004a). Reorientation in a two-dimensional environment: I. Do humans encode the featural and geometric properties of a two-dimensional schematic of a room? Journal of Comparative Psychology, 118, 82-94. Abstract
  • MacDonald, S.E., Spetch, M.L., Kelly, D.M & Cheng, K. (2004). Strategies for landmark use by children, adults and marmoset monkeys. Learning & Motivation, 35, 322-347. Abstract
  • Gray, E.R., Spetch, M.L., Kelly, D.M., & Nguyen, A. (2004). Searching in the center: pigeons encode relative distances from the walls of an enclosure. Journal of Comparative Psychology, 118, 113-117. DOI: 10.1037/0735-7036.118.1.113 Abstract


  • Cook, R.G., Kelly, D.M. & Katz, J.S. (2003). Successive two-item same-different discrimination and concept learning by pigeons. Behavioral Processes, 62, 125-144. Abstract
  • Kelly, D.M. & Cook, R.G. (2003). Differential effects of visual context on pattern discrimination by pigeons and humans. Journal of Comparative Psychology, 117, 200-208. Abstract


  • Kelly, D.M. & Spetch, M.L (2001). Pigeons encode relative geometry. Journal of Experimental Psychology: Animal Behavior Processes, 27, 417-422. Abstract
  • Kelly, D.M., Bischof, W.F., Wong-Wylie, D.R. & Spetch, M.L. (2001). Detection of Glass patterns by pigeons and humans: implications for differences in higher-level processing. Psychological Science, 12, 338-342. DOI: 10.1111/1467-9280.00362 Abstract


  • Spetch, M.L., Kelly, D.M. & Reid, S. (1999). Recognition of objects and spatial relations in pictures across changes in viewpoint. Current Psychology of Cognition, 18, 729-764. Full Article
  • Cook, R.G., Katz, J.S., & Kelly, D.M. (1999). Pictorial same-different concept learning and discrimination in pigeons. Current Psychology of Cognition, 18, 805-843.


  • Kelly, D.M., Spetch, M.L. & Heth, C.D. (1998) Pigeons' encoding of geometric and featural properties of a spatial environment. Journal of Comparative Psychology, 112,259-269.
  • Spetch, M.L., Kelly, D.M. & Lechelt, D.P. (1998). Encoding of spatial information in images of an outdoor scene by pigeons and humans. Animal Learning & Behavior, 26,85-102. Full Article


  • Spetch, M.L. Cheng, K., MacDonald, S.E., Linkenhoker, B.A., Kelly, D.M. & Doerkson, S.R. (1997). Use of landmark configuration in pigeons and humans: II. Generality across search tasks. Journal of Comparative Psychology, 111, 14-24. Abstract

Book Chapters

  • Reichert, J.F., Schwarz, S. & Kelly, D.M. (in submission). Spatial cognition in birds. In: Healy, S. and Ten Cate, C., (Eds.). Avian Cognition.
  • Wilzeck, C. & Kelly, D.M. (2012). Avian visual pseudoneglect: The effect of age and sex on visuospatial side biases. Behavioural lateralization in vertebrates – two sides of the same coin. Springer-Verlag.
  • Kelly, D.M. (2012). Animal Learning. Oxford Bibliographies in Psychology. Ed. D.S. Dunn. New York: Oxford University Press,
  • Kelly, D.M. & Spetch, M.L. (2010). Comparative Spatial Cognition: Encoding of geometric information from surfaces and landmark arrays. In Wasserman and Zentall (Ed). Comparative Cognition: Experimental explorations into animal intelligence. (pp. 210-228). Oxford: Oxford University Press.
  • Spetch, M.L. & Kelly, D.M. (2006). Comparative Spatial Cognition: Processes in landmark and surface-based place finding. In Wasserman and Zentall (Ed). Comparative Cognition: Experimental explorations into animal intelligence. (pp. 210-228). Oxford: Oxford University Press.
  • Spetch, M.L., Kelly, D.M. & Reid, S. (2000). Recognition of objects and spatial relations in pictures across changes in viewpoint. In J. Fagot (Ed). Picture perception in animals(pp.107-142). Philadelphia. PA: Psychology Press.
  • Cook, R.G., Katz, J.S., & Kelly, D.M. (2000). Pictorial same-different concept learning and discrimination in pigeons. In J. Fagot (Ed). Picture perception in animals (pp.181-218). Philadelphia. PA: Psychology Press.