#
# Keys (Column 3: "type") constrained to be either:
# (a) a term from the "lite" sequence ontology, SOFA;
# (b) a SOFA accession number (SO:000000). 
#
# key               Qualifiers (@=mandatory)
# =======           ========================
centromere          ID Name Alias Parent Note Target Gap Derives_from feature_id isObsolete timelastmodified note comment private curation
region              ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term Start_range End_range feature_id isObsolete timelastmodified description locus stable_id EC_number gene_symbol Ontology_term dbxref molecule_type size organism_name strain topology localization gff_source gff_seqname score 
contig              ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term Start_range End_range feature_id isObsolete timelastmodified description locus stable_id EC_number gene_symbol Ontology_term Dbxref molecule_type size organism_name strain topology localization gff_source gff_seqname score comment
#supercontig         ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified description locus stable_id EC_number gene_symbol Ontology_term Dbxref molecule_type size organism_name strain topology localization gff_source gff_seqname score 
#chromosome          ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified note comment description locus stable_id EC_number gene_symbol Ontology_term Dbxref molecule_type size organism_name strain topology localization gff_source gff_seqname score 
gene                ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term Start_range End_range feature_id isObsolete timelastmodified locus stable_id EC_number gene_symbol Ontology_term Dbxref molecule_type size organism_name strain topology localization gff_source gff_seqname score private comment
#CDS                 ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified codon_start description locus stable_id EC_number gene_symbol Ontology_term Dbxref molecule_type size organism_name strain topology localization gff_source gff_seqname score  results note comment job
polypeptide         ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term Start_range End_range feature_id isObsolete timelastmodified description locus stable_id EC_number gene_symbol Ontology_term Dbxref product molecule_type size organism_name strain topology localization gff_source gff_seqname score mass isoelectric charge source private signal_peptide membrane_structure cytoplasmic_polypeptide_region non_cytoplasmic_polypeptide_region transmembrane_polypeptide_region cytoplasm_location transmembrane non_cytoplasm_location GPI_anchor_cleavage_site comment curation translation
pseudogene          ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term Start_range End_range feature_id isObsolete timelastmodified description locus stable_id EC_number gene_symbol Ontology_term Dbxref molecule_type size organism_name strain topology localization gff_source gff_seqname score private
pseudogenic_exon    ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term Start_range End_range feature_id isObsolete timelastmodified description locus stable_id EC_number gene_symbol Ontology_term Dbxref molecule_type size organism_name strain topology localization gff_source gff_seqname score private
pseudogenic_transcript ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term Start_range End_range feature_id isObsolete timelastmodified description locus stable_id EC_number gene_symbol Ontology_term Dbxref molecule_type size organism_name strain topology localization gff_source gff_seqname score product
intron              ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified description locus stable_id EC_number gene_symbol Ontology_term Dbxref molecule_type size organism_name strain topology localization gff_source gff_seqname score 
exon                ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term Start_range End_range feature_id isObsolete timelastmodified codon_start description locus stable_id EC_number gene_symbol Ontology_term Dbxref molecule_type size organism_name strain topology localization gff_source gff_seqname score 
three_prime_UTR     ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term Start_range End_range feature_id isObsolete timelastmodified systematic_id note comment private curation
five_prime_UTR      ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term Start_range End_range feature_id isObsolete timelastmodified systematic_id note comment private curation
transcript          ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term Start_range End_range feature_id isObsolete timelastmodified private comment
primary_transcript  ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term Start_range End_range feature_id isObsolete timelastmodified product note comment controlled_curation 
tRNA                ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term Start_range End_range feature_id isObsolete timelastmodified product note comment controlled_curation private locus_tag
rRNA                ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term Start_range End_range feature_id isObsolete timelastmodified product note comment controlled_curation private locus_tag
tmRNA                ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term Start_range End_range feature_id isObsolete timelastmodified product note comment controlled_curation private locus_tag
mRNA                ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term Start_range End_range feature_id isObsolete timelastmodified product note comment controlled_curation private locus_tag
ncRNA               ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term Start_range End_range feature_id isObsolete timelastmodified product note comment controlled_curation private locus_tag
snRNA               ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term Start_range End_range feature_id isObsolete timelastmodified product note comment controlled_curation private locus_tag
scRNA               ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term Start_range End_range feature_id isObsolete timelastmodified product note comment controlled_curation private locus_tag
snoRNA              ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term Start_range End_range feature_id isObsolete timelastmodified product note comment controlled_curation private locus_tag
polypeptide_domain  ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified 
polypeptide_motif   ID Name Alias Parent Note Target Gap Derives_from feature_id isObsolete timelastmodified
signal_peptide      ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified 
match               ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified 
nucleotide_match    ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified gff_source gff_seqname score 
translated_nucleotide_match    ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified
protein_match       ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified cluster gff_source gff_seqname score
sequence_difference ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified
direct_repeat       ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified
inverted_repeat     ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified
dispersed_repeat    ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified
repeat_family       ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified gff_source gff_seqname score 
transposable_element  ID Name Alias Parent Target Gap Derives_from Note Dbxref Ontology_term feature_id isObsolete timelastmodified
binding_site        ID Name Alias Parent Target Gap Derives_from Note Dbxref Ontology_term feature_id isObsolete timelastmodified

#minus_10_signal     ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified allele citation evidence function gene label locus_tag map note comment operon partial standard_name 
#minus_35_signal     ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified allele citation evidence function gene label locus_tag map note comment operon partial standard_name 
#three_prime_clip    ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified allele citation evidence function gene label locus_tag map note comment partial standard_name 
#five_prime_clip     ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified allele citation evidence function gene label locus_tag map note comment partial standard_name 
#CAAT_signal         ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified allele citation evidence gene label locus_tag map note comment partial 
#D_loop              ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified allele citation evidence gene label locus_tag map note comment partial 
#GC_signal           ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified allele citation evidence gene label locus_tag map note comment partial 
three_prime_LTR     ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified allele citation evidence function gene label locus_tag map note comment partial standard_name 
five_prime_LTR      ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified allele citation evidence function gene label locus_tag map note comment partial standard_name 
N_region        	ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified allele citation codon codon_start evidence gene label locus_tag map note comment partial product pseudo standard_name 
#STS             	ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified allele citation evidence gene label locus_tag map note comment partial standard_name 
#TATA_box        	ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified allele citation evidence gene label locus_tag map note comment partial 
#attenuator      	ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified allele citation evidence gene label locus_tag map note comment operon partial phenotype 
#enhancer        	ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified allele citation evidence gene label locus_tag map note comment partial standard_name 
gap             	ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified @estimated_length score
#iDNA            	ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified allele citation evidence function gene label locus_tag map note comment number partial standard_name 
#mature_peptide  	ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified EC_number allele citation codon codon_start evidence function gene label locus_tag map note comment partial product pseudo standard_name 
#modified_base_site  ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified @mod_base allele citation evidence frequency gene label locus_tag map note comment 
operon          	ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified @operon allele citation evidence function label map note comment phenotype pseudo standard_name 
polyA_signal_sequence  	ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified allele citation evidence gene label locus_tag map note comment partial 
polyA_site      		ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified allele citation evidence gene label locus_tag map note comment 
#precursor_RNA   		ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified allele citation evidence function gene label locus_tag map note comment operon partial product standard_name 
primary_transcript 		ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified allele citation evidence function gene label locus_tag map note comment operon partial standard_name 
#primer_binding_site     ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified PCR_conditions allele citation evidence gene label locus_tag map note comment partial standard_name 
promoter        		ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified allele citation evidence function gene label locus_tag map note comment operon partial phenotype pseudo standard_name 
#protein_binding_site    ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified @bound_moiety allele citation evidence function gene label locus_tag map note comment partial standard_name 
origin_of_replication	ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified allele citation direction evidence gene label locus_tag map note comment partial standard_name 
repeat_region   	ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified allele citation evidence function gene insertion_seq label locus_tag map note comment partial rpt_family rpt_type rpt_unit standard_name DNA_transposon 
repeat_unit     	ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified allele citation evidence function gene label locus_tag map note comment partial rpt_family rpt_type rpt_unit 
#satellite_DNA       ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified allele citation evidence gene label locus_tag map note comment partial rpt_family rpt_type rpt_unit standard_name 
#stem_loop       	ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified allele citation evidence function gene label locus_tag map note comment operon partial standard_name 
#terminator      	ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified allele citation evidence gene label locus_tag map note comment operon partial standard_name 
#transit_peptide 	ID Name Alias Parent Note Target Gap Derives_from Dbxref Ontology_term feature_id isObsolete timelastmodified allele citation codon codon_start evidence function gene label locus_tag map note comment partial product pseudo standard_name